Basic Science Pathology

Last Minute Review

Pathology — Last Minute Review

Rapid Review

A last-minute review of high-yield neuropathology facts for the RITE and board exams. Tables, key associations, and must-know one-liners — designed for a quick pass the night before.

CNS Tumors
TumorWHO GradeLocationHistology / MarkerKey Fact
Glioblastoma (GBM)4Cerebral hemispheres (adults)Pseudopalisading necrosis, microvascular proliferation; GFAP+; IDH-wildtypeMost common primary malignant brain tumor in adults; “butterfly” pattern crosses corpus callosum
Astrocytoma, IDH-mutant2–4Cerebral hemispheres (young adults)IDH1 R132H mutation; p53 loss; ATRX lossBetter prognosis than IDH-wildtype; 2-HG oncometabolite produced
Oligodendroglioma2–3Frontal lobe (adults)“Fried egg” cells, chicken-wire vasculature; 1p/19q codeletion + IDH-mutantCalcifications on imaging; chemosensitive (PCV); seizures common
Ependymoma2–34th ventricle (children), spinal cord (adults)Perivascular pseudorosettes, true rosettes; GFAP+Posterior fossa → fills 4th ventricle; ZFTA fusion in supratentorial
Medulloblastoma4Posterior fossa (cerebellum, children)Homer Wright rosettes; synaptophysin+Most common malignant pediatric brain tumor; WNT subtype → best prognosis; SHH subtype → PTCH1 mutation (Gorlin)
Meningioma1–3Dura (convexity, parasagittal, sphenoid wing)Psammoma bodies, whorls; EMA+; loss of NF2/merlinMost common primary intracranial tumor overall; extra-axial, dural tail sign
Schwannoma1CN VIII (vestibular, CPA); spinal nerve rootsVerocay bodies, Antoni A (compact) & B (loose); S100+Bilateral vestibular schwannomas → NF2; benign, encapsulated
Craniopharyngioma (Adamantinomatous)1Suprasellar (children & adolescents)Wet keratin, calcifications, cholesterol crystals; β-catenin mutation (CTNNB1)Bimodal age (5–14, 50–74); “machinery oil” cyst fluid; hypothalamic/pituitary dysfunction
Craniopharyngioma (Papillary)1Suprasellar / 3rd ventricle (adults)Well-differentiated squamous epithelium; BRAF V600E mutationAdults only; no calcification; solid; responds to BRAF inhibitors
Pituitary Adenoma1Sella turcicaMonomorphic cells; specific hormone stains; no reticulin networkMost common sellar tumor; prolactinoma most common functional type; bitemporal hemianopia from optic chiasm compression
Hemangioblastoma1Posterior fossa (cerebellar), spinal cordStromal cells with lipid vacuoles, rich capillary networkAssociated with VHL syndrome (chromosome 3p); may cause secondary polycythemia (EPO production)
PCNSLHigh-gradePeriventricular white matter (deep structures)Diffuse large B-cell lymphoma; CD20+; perivascular cuffingImmunocompromised (HIV/EBV) or elderly immunocompetent; “ghost tumor” — shrinks with steroids; do NOT biopsy after steroids
DNET (Dysembryoplastic Neuroepithelial Tumor)1Temporal lobe cortex (children/young adults)Specific glioneuronal element, floating neurons in mucin poolsChronic epilepsy; cortical-based, bubbly/multicystic on MRI; excellent prognosis
Ganglioglioma1–3Temporal lobe (children/young adults)Ganglion cells + glial cells; CD34+; BRAF V600E commonMost common tumor causing chronic epilepsy in children; cystic + mural nodule
Pilocytic Astrocytoma1Posterior fossa / cerebellum (children); optic pathwayRosenthal fibers, eosinophilic granular bodies; BRAF-KIAA1549 fusionMost common pediatric brain tumor; cystic + enhancing mural nodule; optic pathway glioma → NF1
Choroid Plexus Papilloma1Lateral ventricle (children), 4th ventricle (adults)Papillary fronds, single-layer cuboidal epitheliumOverproduction of CSF → hydrocephalus; transthyretin+
Pineoblastoma4Pineal region (children)Small round blue cells; synaptophysin+Trilateral retinoblastoma = bilateral retinoblastoma + pineoblastoma (RB1 loss)
Pineocytoma1Pineal region (adults)Pineocytomatous rosettes; well-differentiatedBenign; Parinaud syndrome (dorsal midbrain compression)
ChordomaLocally aggressiveClivus / sacrum (midline)Physaliphorous cells (bubbly, vacuolated); brachyury+Arises from notochord remnants; locally destructive; high recurrence
Colloid CystBenignAnterior 3rd ventricle (foramen of Monro)Simple cuboidal/columnar epithelium; mucin-filledPositional headaches; acute obstructive hydrocephalus → sudden death risk
Brain MetastasesN/AGray-white junction (hematogenous)Resembles primary tumor; cytokeratin/TTF-1/etc.Most common brain tumors overall; lung > breast > melanoma > renal > colon; melanoma & renal → hemorrhagic mets
💎 Board Pearl
  • IDH-mutant = better prognosis in gliomas — always check IDH status first
  • 1p/19q codeletion is diagnostic of oligodendroglioma (must have both codeletion + IDH mutation)
  • WHO 2021 classification: molecular markers trump histology for diagnosis
  • Butterfly GBM: tumor crossing the corpus callosum — classic imaging finding
  • MGMT promoter methylation → better temozolomide response in GBM
Neurocutaneous Syndromes (Phakomatoses)
SyndromeGeneChromosomeInheritanceKey FeaturesAssociated Tumors
NF1 (von Recklinghausen)NF1 (neurofibromin)17q11AD≥6 café-au-lait macules, axillary freckling, Lisch nodules, bony dysplasiaNeurofibromas (plexiform), optic pathway glioma, MPNST, pheochromocytoma
NF2NF2 (merlin/schwannomin)22q12ADBilateral vestibular schwannomas (hallmark), cataracts, hearing lossSchwannomas, meningiomas, ependymomas
Tuberous Sclerosis (TSC)TSC1 (hamartin) / TSC2 (tuberin)9q34 / 16p13ADCortical tubers, ash-leaf spots, shagreen patch, facial angiofibromas, seizures, intellectual disabilitySubependymal giant cell astrocytoma (SEGA), cardiac rhabdomyoma, renal angiomyolipoma
VHLVHL3p25ADHemangioblastomas (CNS/retina), renal cysts, pheochromocytoma, secondary polycythemiaHemangioblastomas, clear cell renal cell carcinoma, pheochromocytoma
Sturge-WeberGNAQ (somatic)9q21Sporadic (not inherited)Port-wine stain (V1), leptomeningeal angiomatosis, tram-track calcifications, seizures, glaucomaNo associated tumors; vascular malformation
Ataxia-TelangiectasiaATM11q22ARCerebellar ataxia, oculocutaneous telangiectasias, immunodeficiency, radiosensitivity, ↑ AFP↑ risk lymphoma & leukemia; DNA repair defect
Cowden SyndromePTEN10q23ADMucocutaneous lesions, macrocephaly, Lhermitte-Duclos (dysplastic gangliocytoma of cerebellum)Breast, thyroid, endometrial carcinomas
Gorlin Syndrome (Basal Cell Nevus)PTCH19q22ADMultiple basal cell carcinomas, jaw keratocysts, calcified falx cerebri, skeletal anomaliesMedulloblastoma (desmoplastic/SHH subtype), basal cell carcinomas
💎 Board Pearl
  • NF1 = chromosome 17 (“17 letters in von Recklinghausen”); NF2 = chromosome 22 (“NF2 → 22”)
  • Sturge-Weber is the only phakomatosis that is NOT inherited — somatic GNAQ mutation
  • TSC: mTOR pathway → treated with everolimus (mTOR inhibitor) for SEGA
  • Ataxia-telangiectasia: ↑ AFP + ↓ IgA + radiosensitivity = classic triad
Metabolic & Storage Diseases
DiseaseEnzyme DeficiencySubstrate AccumulatedInheritanceKey Features
Tay-SachsHexosaminidase AGM2 gangliosideARCherry-red spot, startle response, progressive neurodegeneration, death by age 3–5; Ashkenazi Jewish
SandhoffHexosaminidase A & BGM2 ganglioside + globosideARLike Tay-Sachs + visceral involvement (hepatosplenomegaly)
Niemann-Pick ASphingomyelinase (acid)SphingomyelinARCherry-red spot, hepatosplenomegaly, “foam cells,” fatal by age 3
Niemann-Pick CNPC1/NPC2 (cholesterol transport)Cholesterol / sphingolipidsARVertical supranuclear gaze palsy, ataxia, dystonia, dementia; juvenile/adult onset possible
Gaucher (Type 1)β-GlucocerebrosidaseGlucocerebrosideAR“Crinkled paper” macrophages, hepatosplenomegaly, bone crises; no CNS involvement in type 1; most common lysosomal storage disease
Gaucher (Types 2 & 3)β-GlucocerebrosidaseGlucocerebrosideARType 2: acute neuropathic, fatal in infancy; Type 3: chronic neuropathic, seizures, eye movement abnormalities
Fabryα-Galactosidase AGlobotriaosylceramide (Gb3)X-linked recessivePainful acroparesthesias, angiokeratomas, corneal verticillata, renal failure, stroke (young male); only X-linked lipidosis
KrabbeGalactocerebrosidaseGalactocerebroside / psychosineARGloboid cells (multinucleated macrophages), peripheral neuropathy, severe demyelination, irritability, death by age 2
Metachromatic Leukodystrophy (MLD)Arylsulfatase ASulfatideARMetachromatic granules (brown with cresyl violet), diffuse demyelination, peripheral neuropathy, gait disturbance
Hurler (MPS I)α-L-IduronidaseHeparan & dermatan sulfateARCoarse facies, corneal clouding, hepatosplenomegaly, dysostosis multiplex, intellectual disability; most severe MPS
Hunter (MPS II)Iduronate-2-sulfataseHeparan & dermatan sulfateX-linked recessiveLike Hurler but NO corneal clouding; only X-linked MPS
Pompe (GSD II)Acid maltase (α-glucosidase)GlycogenARCardiomegaly (infantile), proximal myopathy, respiratory failure; ERT available (alglucosidase alfa)
GM1 Gangliosidosisβ-GalactosidaseGM1 gangliosideARCherry-red spot, hepatosplenomegaly, coarse facies, skeletal changes; resembles Hurler + Tay-Sachs combined
Neuronal Ceroid Lipofuscinosis (NCL/Batten)Various (CLN1–CLN14)Lipofuscin (autofluorescent)ARSeizures, vision loss, progressive dementia; “curvilinear bodies” or “fingerprint profiles” on EM
💎 Board Pearl
  • Cherry-red spot: Tay-Sachs, Niemann-Pick A, Sandhoff, GM1 gangliosidosis — ganglion cells full of lipid except at fovea
  • Fabry & Hunter = X-linked (“Fabulous Hunters are X-linked”)
  • Gaucher is the most common lysosomal storage disease; GBA mutations → ↑ risk of Parkinson disease
  • Pompe = only glycogen storage disease that is also a lysosomal storage disease
Leukodystrophies
DiseaseGene / EnzymeInheritanceMRI PatternKey Feature
Metachromatic Leukodystrophy (MLD)Arylsulfatase A (ARSA gene)ARConfluent periventricular white matter → frontal predominant; spares subcortical U-fibers earlyMetachromatic granules on nerve biopsy; peripheral neuropathy + central demyelination
Krabbe (Globoid Cell)Galactocerebrosidase (GALC gene)ARPeriventricular + cerebellar white matter; thalamic hyperdensity on CTGloboid cells; extreme irritability in infancy; CSF protein elevated; psychosine toxicity
X-linked Adrenoleukodystrophy (X-ALD)ABCD1 (peroxisomal VLCFA transport)X-linked recessivePosterior periventricular → splenium first; enhancing leading edge of demyelination↑ VLCFA; boys: cerebral form; adults: adrenomyeloneuropathy (spastic paraparesis); adrenal insufficiency
Canavan DiseaseAspartoacylase (ASPA gene)ARDiffuse white matter + subcortical U-fibers involved; macrocephaly↑ NAA on MRS and urine; Ashkenazi Jewish; spongiform degeneration; macrocephaly
Alexander DiseaseGFAP (gain-of-function)AD (de novo)Frontal predominant; periventricular rim; basal ganglia/thalamic involvementRosenthal fibers (GFAP aggregates); macrocephaly; seizures; frontal → posterior progression
Pelizaeus-Merzbacher Disease (PMD)PLP1 (proteolipid protein 1)X-linked recessiveDiffuse T2 hyperintensity of white matter (“tigroid” pattern); lack of myelinationNystagmus from birth; connatal form severe; hypomyelinating leukodystrophy
Vanishing White Matter DiseaseeIF2B subunits (EIF2B1–5)ARDiffuse white matter → cystic rarefaction (CSF signal replacement)Episodic deterioration with febrile illness or minor head trauma; ovarian failure in females (ovarioleukodystrophy)
💎 Board Pearl
  • MRI pattern helps differentiate: frontal-predominant (Alexander, MLD late) vs. posterior-predominant (X-ALD) vs. diffuse + U-fibers (Canavan)
  • Canavan vs. Alexander: both have macrocephaly, but Canavan = ↑ NAA, Alexander = Rosenthal fibers
  • ↑ NAA on MRS = Canavan disease (nearly pathognomonic)
  • X-ALD: Lorenzo’s oil does NOT reverse existing CNS damage — HSCT is the only effective treatment for cerebral form
Mitochondrial Disorders
SyndromeMutationInheritanceKey Features
MELASm.3243A>G (MT-TL1, tRNA-Leu)Maternal (mtDNA)Stroke-like episodes (not vascular territory), seizures, lactic acidosis, short stature, ↑ lactate; ragged red fibers
MERRFm.8344A>G (MT-TK, tRNA-Lys)Maternal (mtDNA)Myoclonus epilepsy, ataxia, ragged red fibers, lipomas; “myoclonus epilepsy with ragged red fibers”
LHONm.11778G>A (most common), m.3460, m.14484Maternal (mtDNA)Bilateral painless subacute vision loss (young males), centrocecal scotoma, pseudoedema of disc; no ragged red fibers
Kearns-SayreLarge-scale mtDNA deletion (>1 kb)Sporadic (de novo deletion)Onset <20 years, progressive external ophthalmoplegia (PEO), pigmentary retinopathy, cardiac conduction block, ↑ CSF protein
NARPm.8993T>G or T>C (MT-ATP6)Maternal (mtDNA)Neuropathy, ataxia, retinitis pigmentosa; at high heteroplasmy (>90%) → Leigh syndrome
Leigh SyndromeVarious (nuclear & mtDNA)Variable (AR, X-linked, maternal)Bilateral symmetric basal ganglia/brainstem lesions; lactic acidosis; developmental regression; most common mitochondrial disease in children
CPEOmtDNA deletions or point mutations (various)Sporadic or maternal or AD/AR (nuclear genes: POLG, TWNK)Progressive ptosis + ophthalmoplegia; ragged red fibers on biopsy; may overlap with Kearns-Sayre
💎 Board Pearl
  • MELAS strokes do NOT respect vascular territories — key differentiator from embolic stroke
  • Ragged red fibers (Gomori trichrome) = accumulation of abnormal mitochondria; present in MELAS, MERRF, KSS, CPEO but NOT LHON
  • Lactic acidosis is universal in mitochondrial disorders — check serum & CSF lactate
  • Kearns-Sayre triad: PEO + retinopathy + cardiac block (onset <20 years)
  • Leigh syndrome MRI: bilateral symmetric T2 hyperintensity in basal ganglia and/or brainstem
Embryology & Developmental Malformations
TimingProcessMalformationKey Feature
Weeks 3–4Primary neurulation (neural tube closure)Anencephaly (rostral closure failure)Absent brain/calvarium above orbits; ↑ AFP; incompatible with life
Weeks 3–4Primary neurulationMyelomeningocele (caudal closure failure)Spina bifida aperta; ↑ AFP; associated with Chiari II; folate prevents
Weeks 3–4Primary neurulationEncephaloceleBrain tissue herniates through skull defect; occipital most common (Western); frontal (Southeast Asian)
Week 5+Secondary neurulationSpinal dysraphism (occult)Tethered cord, lipomyelomeningocele, dermal sinus; skin markers (dimple, tuft, hemangioma)
Weeks 5–6Prosencephalic development (cleavage)HoloprosencephalyFailed forebrain cleavage; alobar (most severe) → semilobar → lobar; SHH, ZIC2, SIX3 genes; trisomy 13
Weeks 5–6Prosencephalic developmentSepto-optic dysplasiaAbsent septum pellucidum, optic nerve hypoplasia, hypothalamic-pituitary dysfunction; HESX1 gene
Weeks 8–16Neuronal migrationLissencephaly (type 1 / classical)Smooth brain (“agyria”); LIS1/PAFAH1B1 (Miller-Dieker, chr 17) or DCX (X-linked, males)
Weeks 8–16Neuronal migrationLissencephaly (type 2 / cobblestone)Overmigration through breached pial surface; Walker-Warburg, muscle-eye-brain disease; associated with congenital muscular dystrophies
Weeks 8–16Neuronal migrationPeriventricular nodular heterotopiaSubependymal gray matter nodules that failed to migrate; FLNA gene (X-linked dominant, lethal in males); seizures
Weeks 8–16Neuronal migrationSubcortical band heterotopia (“double cortex”)DCX gene (X-linked, females); band of gray matter between cortex and ventricle
Late fetalCortical organizationPolymicrogyria (PMG)Excessive small, fused gyri; perisylvian most common; acquired (CMV infection) or genetic
Weeks 5–6Rhombencephalon developmentDandy-Walker MalformationHypoplastic/absent cerebellar vermis, cystic dilation of 4th ventricle, enlarged posterior fossa
N/AHindbrain/foramen magnumChiari ICerebellar tonsillar herniation ≥5 mm below foramen magnum; syringomyelia; headache with Valsalva
N/AHindbrain/neural tubeChiari IICerebellar tonsillar + vermian herniation; ALWAYS with myelomeningocele; beaked tectum, small posterior fossa
Weeks 8–17Commissural developmentAgenesis of Corpus CallosumColpocephaly (dilated occipital horns), parallel lateral ventricles, “racing car sign” on axial MRI; may be isolated or syndromic
💎 Board Pearl
  • Chiari II is ALWAYS associated with myelomeningocele — Chiari I is NOT
  • Lissencephaly: LIS1 = posterior predominant, DCX = anterior predominant
  • Folate supplementation prevents neural tube defects — start before conception
  • Holoprosencephaly + midline facial defects + trisomy 13 = classic association
  • Periventricular heterotopia (FLNA) → seizures in young females with normal intelligence
Neurogenetics
DiseaseGeneInheritanceRepeat / MutationKey Feature
Huntington DiseaseHTT (huntingtin)ADCAG repeat ≥36 (chr 4p); anticipation (paternal)Chorea, psychiatric symptoms, dementia; caudate atrophy; onset ~40s; juvenile form: rigidity + seizures (Westphal variant)
Friedreich AtaxiaFXN (frataxin)ARGAA repeat expansion (intron 1); loss of frataxinProgressive ataxia, cardiomyopathy, diabetes, pes cavus, scoliosis; dorsal columns + spinocerebellar tracts + peripheral nerves; most common hereditary ataxia
SCA1ATXN1ADCAG repeatCerebellar ataxia + pyramidal signs; brainstem/cerebellar atrophy
SCA2ATXN2ADCAG repeatSlow saccades (hallmark) + ataxia + neuropathy
SCA3 (Machado-Joseph)ATXN3ADCAG repeatMost common SCA worldwide; bulging eyes, ataxia, dystonia, spasticity
SCA6CACNA1AADCAG repeat (small expansion)Pure cerebellar ataxia; allelic with episodic ataxia type 2 and familial hemiplegic migraine
SCA7ATXN7ADCAG repeatAtaxia + retinal degeneration (macular dystrophy) — only SCA with vision loss
Fragile X SyndromeFMR1X-linked dominantCGG repeat ≥200 (full mutation); premutation 55–200Most common inherited cause of intellectual disability; macroorchidism, long face, large ears; premutation carriers → FXTAS (tremor/ataxia in elderly males)
Myotonic Dystrophy Type 1 (DM1)DMPKADCTG repeat (3’ UTR); anticipation (maternal for congenital)Distal weakness, myotonia, cataracts, cardiac conduction, frontal balding, insulin resistance; most common adult muscular dystrophy
Myotonic Dystrophy Type 2 (DM2)CNBP (ZNF9)ADCCTG repeat (intron 1)Proximal weakness (“PROMM”); less severe than DM1; no congenital form; myotonia less prominent
Duchenne Muscular Dystrophy (DMD)DMD (dystrophin)X-linked recessiveFrameshift/nonsense → absent dystrophinOnset <5 yrs; pseudohypertrophy of calves, Gowers sign; CK >10,000; loss of ambulation by ~12 yrs; cardiomyopathy
Becker Muscular Dystrophy (BMD)DMD (dystrophin)X-linked recessiveIn-frame deletion → reduced/truncated dystrophinMilder than DMD; ambulation preserved into teens/adulthood; cardiomyopathy may predominate
Spinal Muscular Atrophy (SMA)SMN1 (survival motor neuron)ARHomozygous deletion of SMN1 (5q13); SMN2 copy number modifies severityLMN disease; Type 1 (Werdnig-Hoffmann) — floppy infant, never sits; Type 2 — sits, never walks; Type 3 (Kugelberg-Welander) — walks
CMT1APMP22ADPMP22 duplication (17p12)Demyelinating; slow NCVs (<38 m/s); pes cavus, distal atrophy, onion bulbs on biopsy; most common CMT
CMT1BMPZ (P0)ADPoint mutationsDemyelinating; similar to CMT1A but earlier onset and more severe
CMT2AMFN2 (mitofusin 2)ADPoint mutationsAxonal; normal NCVs; reduced CMAP amplitudes; most common axonal CMT
HNPPPMP22ADPMP22 deletion (opposite of CMT1A)Episodic, painless, pressure-sensitive palsies; tomaculae (“sausage-shaped” myelin thickenings) on biopsy
Wilson DiseaseATP7BARVarious point mutations (chr 13)Copper accumulation → Kayser-Fleischer rings, dystonia, parkinsonism, psychiatric, liver disease; ↓ ceruloplasmin, ↑ 24h urine copper
Familial ALS (SOD1)SOD1ADPoint mutations (most common: A4V — aggressive)~2% of all ALS; posterior column involvement may occur (unlike sporadic); copper-zinc superoxide dismutase
Familial ALS (C9orf72)C9orf72ADGGGGCC hexanucleotide repeat expansionMost common genetic cause of ALS and FTD; ALS-FTD spectrum; RNA foci + dipeptide repeat proteins
💎 Board Pearl
  • Trinucleotide repeat + anticipation = Huntington (CAG), DM1 (CTG), Fragile X (CGG), Friedreich (GAA), SCAs (CAG)
  • CMT1A = PMP22 duplication; HNPP = PMP22 deletion — mirror disorders of the same gene
  • DM1 vs DM2: DM1 = distal, DM2 = proximal; DM1 has congenital form, DM2 does not
  • C9orf72 = most common genetic cause of BOTH familial ALS and familial FTD
  • SMA: SMN2 copy number determines severity — nusinersen, onasemnogene, risdiplam all target SMN2
Neuropathology Buzzwords
FindingDiagnosis
Rosenthal fibersPilocytic astrocytoma; Alexander disease
Eosinophilic granular bodies (EGBs)Pilocytic astrocytoma; ganglioglioma; pleomorphic xanthoastrocytoma
“Fried egg” cellsOligodendroglioma (artifact of formalin fixation)
Chicken-wire capillary patternOligodendroglioma
Pseudopalisading necrosisGlioblastoma (GBM)
Psammoma bodiesMeningioma; papillary thyroid carcinoma; serous ovarian carcinoma
WhorlsMeningioma
Homer Wright rosettesMedulloblastoma; neuroblastoma; pineoblastoma
Perivascular pseudorosettesEpendymoma
True ependymal rosettesEpendymoma (rare but specific)
Verocay bodiesSchwannoma
Antoni A (compact, palisading) & Antoni B (loose, myxoid)Schwannoma
Physaliphorous cells (bubbly vacuolated cytoplasm)Chordoma
Cherry-red spot (macula)Tay-Sachs, Niemann-Pick A, Sandhoff, GM1 gangliosidosis; also central retinal artery occlusion
Crinkled/wrinkled paper macrophagesGaucher disease
Foam cellsNiemann-Pick disease
Globoid cells (multinucleated macrophages)Krabbe disease
Metachromatic granules (brown with cresyl violet)Metachromatic leukodystrophy
Negri bodies (intracytoplasmic eosinophilic inclusions)Rabies
Cowdry type A inclusions (intranuclear eosinophilic with halo)HSV encephalitis; VZV; CMV
Cowdry type B inclusions (ground-glass intranuclear)Poliomyelitis; other enteroviruses
Owl-eye inclusionsCMV (cytomegalovirus)
Lewy bodies (intracytoplasmic, eosinophilic, halo)Parkinson disease; Lewy body dementia
Neurofibrillary tangles (hyperphosphorylated tau)Alzheimer disease; CTE; PSP; other tauopathies
Amyloid plaques (Aβ42)Alzheimer disease
Pick bodies (round, intracytoplasmic tau inclusions)Pick disease (bvFTD variant)
Hirano bodies (rod-shaped, eosinophilic, actin-rich)Alzheimer disease (hippocampus); also normal aging
Granulovacuolar degenerationAlzheimer disease (hippocampal neurons)
Bunina bodies (small eosinophilic, intracytoplasmic)ALS (motor neurons)
TDP-43 inclusions (ubiquitinated, cytoplasmic)ALS; frontotemporal dementia (FTD-TDP)
Ragged red fibers (Gomori trichrome)Mitochondrial myopathies (MELAS, MERRF, KSS)
Onion bulbs (Schwann cell concentric layers)Chronic demyelinating neuropathy (CMT1A, CIDP)
Tomaculae (“sausage-shaped” myelin swellings)HNPP (hereditary neuropathy with pressure palsies)
Zebra bodies (lamellar inclusions on EM)Fabry disease; other storage diseases
Curvilinear profiles / fingerprint bodies (EM)Neuronal ceroid lipofuscinosis (NCL/Batten)
Clinical Pearl

If a board question describes a histological finding, match it to the diagnosis using this table. These associations are tested repeatedly. “Pseudopalisading necrosis” = GBM and “fried egg cells” = oligodendroglioma are the two most commonly tested neuropathology buzzwords.

Neurotoxicology & Nutritional Deficiencies
Toxin / DeficiencyKey Neurological Finding
Lead (Pb)Children: encephalopathy, cognitive decline, ↓ IQ, lead lines on gingiva, basophilic stippling; Adults: peripheral neuropathy (wrist/foot drop), abdominal colic; ↑ free erythrocyte protoporphyrin
Mercury (organic/methyl)Visual field constriction, ataxia, paresthesias, cognitive decline; Minamata disease; prenatal → cerebral palsy
ArsenicPainful sensorimotor axonal neuropathy, Mees lines (white transverse nail lines), GI symptoms, “rice water” diarrhea, skin changes
ManganeseParkinsonism (“manganism”) — gait abnormality, dystonia, psychiatric; globus pallidus T1 hyperintensity on MRI; does NOT respond to levodopa
Carbon monoxide (CO)Acute: headache, confusion, cherry-red skin; Delayed: parkinsonism, cognitive decline; globus pallidus necrosis; white matter demyelination
ThalliumPainful sensory neuropathy, alopecia, Mees lines; rat poison
OrganophosphatesCholinergic crisis (SLUDGE: salivation, lacrimation, urination, defecation, GI distress, emesis); intermediate syndrome (proximal weakness days later); delayed polyneuropathy
MethanolVisual loss (retinal toxicity → optic nerve), metabolic acidosis with ↑ osmolar gap; putaminal necrosis
Vitamin B1 (thiamine) deficiencyWernicke encephalopathy: confusion, ophthalmoplegia, ataxia (classic triad); mammillary body/periaqueductal hemorrhage; Korsakoff: anterograde amnesia, confabulation
Vitamin B6 (pyridoxine) deficiencyPeripheral neuropathy, seizures (neonates); isoniazid-induced; EXCESS B6 also causes sensory neuropathy
Vitamin B12 (cobalamin) deficiencySubacute combined degeneration (dorsal columns + lateral corticospinal tracts), megaloblastic anemia, cognitive decline; ↑ methylmalonic acid, ↑ homocysteine
Vitamin E (α-tocopherol) deficiencySpinocerebellar syndrome resembling Friedreich ataxia; ataxia, proprioceptive loss, areflexia; fat malabsorption, abetalipoproteinemia
Copper deficiencyMyelopathy mimicking B12 deficiency (subacute combined degeneration); anemia, neutropenia; zinc excess or gastric surgery causes copper depletion
Wernicke EncephalopathyThiamine deficiency; give thiamine BEFORE glucose; mammillary bodies, medial thalami, periaqueductal gray, tectal plate
Marchiafava-Bignami DiseaseCorpus callosum demyelination/necrosis; associated with chronic alcoholism and nutritional deficiency; acute: coma/seizures; chronic: dementia, gait abnormalities
💎 Board Pearl
  • Always give thiamine BEFORE glucose in suspected Wernicke — glucose without thiamine worsens the condition
  • B12 deficiency vs. copper deficiency: clinically identical myelopathy — check copper if B12 is normal
  • Manganese: T1 hyperintensity in globus pallidus (not T2) — unique MRI finding
  • Lead in children = encephalopathy; lead in adults = neuropathy
  • Vitamin E deficiency mimics Friedreich ataxia — treatable cause, always check levels
Classic Board Traps

Board Traps — Do Not Get Fooled

  • Meningioma vs. metastasis: Both are extra-axial and enhance, but meningioma has dural tail, broad dural base, and calcifications; metastasis has no dural tail and is often multiple
  • Oligodendroglioma vs. astrocytoma: Both are infiltrative gliomas, but oligodendroglioma requires 1p/19q codeletion + IDH mutation — histology alone is insufficient under WHO 2021
  • Pilocytic astrocytoma vs. GBM: Both can enhance, but pilocytic is grade 1 (cystic + mural nodule, children, posterior fossa) while GBM is grade 4 (ring-enhancing, necrosis, adults, hemispheric)
  • PCNSL vs. GBM: Both are deep hemispheric lesions, but PCNSL → homogeneous enhancement, periventricular, responds to steroids (“ghost tumor”); GBM → ring enhancement with necrosis
  • NF1 vs. NF2: NF1 = neurofibromas + optic glioma + café-au-lait spots (chromosome 17); NF2 = schwannomas + meningiomas (chromosome 22). Do NOT confuse the tumor types
  • Tay-Sachs vs. Niemann-Pick A: Both have cherry-red spot, but only Niemann-Pick has hepatosplenomegaly (Tay-Sachs = NO visceral involvement)
  • Canavan vs. Alexander: Both cause macrocephaly in infants with leukodystrophy, but Canavan = ↑ NAA (MRS) and Alexander = Rosenthal fibers + frontal predominance on MRI
  • X-ALD MRI pattern: Posterior white matter → anterior (parieto-occipital); Alexander is the opposite (frontal → posterior)
  • MELAS strokes: Do NOT follow vascular territories — if a young patient has “stroke” that crosses vascular boundaries + lactic acidosis, think MELAS, not embolic stroke
  • Friedreich ataxia: AR, NOT AD — do not confuse with the SCAs which are AD; also Friedreich = GAA repeat (not CAG)
  • CMT1A vs. HNPP: Both involve PMP22 on chromosome 17, but CMT1A = duplication (too much PMP22) and HNPP = deletion (too little PMP22)
  • DM1 vs. DM2: DM1 = distal weakness + congenital form possible; DM2 = proximal weakness + NO congenital form
  • Copper deficiency vs. B12 deficiency: Identical myelopathy (subacute combined degeneration) — if B12 is normal, check copper before diagnosing idiopathic myelopathy
  • Manganese parkinsonism: T1 hyperintensity (not T2) in globus pallidus; does NOT respond to levodopa — distinguishes from idiopathic Parkinson disease
  • Chiari I vs. Chiari II: Chiari I = tonsillar herniation only, NO myelomeningocele; Chiari II = ALWAYS with myelomeningocele + more extensive herniation
  • Lissencephaly genetics: LIS1 (17p) = posterior-predominant smooth brain; DCX (Xq) = anterior-predominant (males) or subcortical band heterotopia (females)
  • Kearns-Sayre vs. CPEO: Both have PEO, but Kearns-Sayre adds pigmentary retinopathy + cardiac conduction block + onset before age 20